This guide shows the locations of all owl towers. The maps found in the library is not complete.
– You cannot get the last rune for the hammer as you need the owl bomb. The quest for the owl bomb cannot be progressed past the underground gate that needs a lever, as the lever you get after fixing at the blacksmith doesn’t fit. If someone found the solution, please let me know as the sub-quests for this is marked completed already.
– One of the runes for the Shield you get after completing a side-quest revolving the sisters who sell the shields. The other three are in towers. You cannot get the shield either, because the NPC sister that gives the shield does not respond even with 4 runes
– Muccia forest has two towers, one when you visit via bobo forest through its west exit, and the other when you visit it via the forgotten path.
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